Cuarta semana, Mesodermo- Capitulo 8, Embriologia Humana Integrada.

[Música]

Buen día. Hoy vamos a conversar acerca de la cuarta semana del desarrollo, especialmente sobre los derivados de la capa mesodérmica. Como siempre, recomendando la lectura de su libro Embriología Humana Integrada, en su segunda edición, en esta ocasión el capítulo octavo.

Entonces, digamos que para este video, pues el mesodermo va a tener una continuidad entre la tercera y la cuarta semana. Pero para, digamos, las explicaciones, vamos a hacer referencia a los eventos que ocurren especialmente en la cuarta semana del desarrollo. Y en este caso, estas son imágenes que nos muestran cortes transversales del disco germinativo trilaminar, en este caso en la cuarta semana, convirtiéndose en una estructura cilíndrica y además en forma de "C". Todo lo que estamos viendo aquí va a estar en medio de la cavidad coriónica, que sería la que estaría por fuera, aquí arriba la cavidad amniótica. Digamos que esto sería la dorsal central. Tendremos el saco vitelino. La línea azul es el ectodermo, la línea amarilla los derivados endodérmicos y para este caso, la línea roja va a ser el mesodermo. Recordando de que vamos a tener unos importantes cambios durante la cuarta semana sobre el mesodermo, secundarios a lo que es el plegamiento transversal.

Entonces, de este mesodermo nosotros vamos a tener cuatro tipos de mesodermo. El primero, digamos que vamos a mencionar, es el que está en la línea media, justo ventral al tubo neural. En este caso, lo denominaremos el notocorda. Y está en la línea media. En principio, es una importante estructura que nos va a producir una izquierda y una derecha, y además son cientos de regulación molecular trascendental en los desarrollos de la tercera y cuarta semana. Sin embargo, en la medida en que se produce la evolución del embrión, va a tener cada vez menos trascendencia en los derivados finales, y lo que nos generará serán los núcleos pulposos de los discos intervertebrales.

Al lado de la línea media, vamos a tener el mesodermo paraaxial, o sea, que está al lado de la axis, o sea, al lado de la línea media. Entonces, este mesodermo paraaxial nos va a generar los somitas. De estos somitas, que van a generar múltiples estructuras, digamos, en principio de sostén, vamos a tener desarrollo de huesos, ligamentos, músculos, dermis. Luego, al lado de él... y ahora que hacer referencia de que este mesodermo paraaxial, vamos a tener derecho e izquierdo. Igual pasa con el mesodermo intermedio y el mesodermo de la placa lateral.

Del mesodermo intermedio, nos va a generar, entre otras estructuras, riñón, parte de las vías urinarias, genitales internos que pueden ser masculinos o femeninos, y también las gónadas. Además de esto, pues vamos a tener el mesodermo de la placa lateral. Este mesodermo nos va a ayudar a conformar, porque va a hacer las paredes del tracto, las paredes musculares del tracto intestinal, respiratorio, en lo que va a ser el mesodermo esplénico, a través del mesodermo de la capa lateral, el mesodermo somático, el cual nos va a ayudar a conformar parte de lo que van a ser las paredes torácica y abdominal, entre otras estructuras importantes.

Entonces, el desarrollo del mesodermo en esta tercera, cuarta semana del desarrollo, van a ser muy, muy, muy particulares, porque nos van a mostrar cómo hay una regulación molecular para que se nos puedan presentar todas estas especificaciones de este tejido celular. Entonces, hay un reloj molecular en donde debe haber una expresión de moléculas, la cual debe ser cíclica y además en momentos específicos, y también debe ser controlada en momentos al alza, en otros a la baja, entre otras moléculas que participan en otros procesos, digamos, dramáticamente importantes para hacer el noche. Toda la vida, toda la vida del noche, dónde y los genes Hox.

Además, digamos que en este caso, hablando un poco particularmente de lo que van a ser los somitas, estos deben aparecer ordenadamente y para cada día del desarrollo. Por ejemplo, el día 21, 22, 23, va a haber una cantidad particular de estos, de estos somitas en ubicaciones especiales. Entonces, digamos que está escrito para que esto ocurra en cada una de estas, digamos, regiones de, en principio, mesodermo paraaxial. Vamos a tener que se elevan las concentraciones y cantidades de Hox. Esto va a generar que se produzca un somita. Luego baja el Hox, y esta somita ya va a estar particularmente diferenciada.

Además, para que haya unos límites claros en estos somitas, pues vamos a tener que en la parte craneal del somita, vamos a tener importantes niveles de ácido retinoico, y en la porción caudal de este, desde ese somita en particular, por supuesto, el ejemplo es para todos, se necesitan niveles de factor de crecimiento fibroblástico tipo 8 y niveles de Wnt-3a.

Veintidós somitas, digamos, van a aparecer, digamos, en días particulares. Por ejemplo, aquí tenemos la tabla claramente. En el día 20, vamos a tener entre 1 y 4. En el 25, entre 17 y 20. Alrededor del día 30, entre 30 y 35. Habría que aclarar que los somitas son entre 42 y 44. Que vamos a tener unas somitas occipitales que van a, por poner un ejemplo, a tener que ver con el desarrollo del cráneo. Vamos a tener unas somitas cervicales, 8. Va a tener somitas torácicos, 12. Cinco lumbares. Vamos a tener los somitas sacrocoxígeos, que son alrededor de 10 a 12.

Entonces, estos somitas se van a tener que ver directamente con el número de vértebras. Y digamos, en los somitas sacrocoxígeos, estos se van a fusionar y al final, pues vamos a tener la estructura específica que sería el sacro y el coxis. Entonces, por ejemplo, nosotros tenemos aquí que hay el día 30, 30, 5. Pero he dicho que en total hay entre 42 y 44. Esto hace referencia a que, en la medida en que aparecen unos, otros pueden ir desapareciendo, y todo esto obedece a la regulación molecular.

Entonces, este mesodermo paraaxial, nosotros lo podemos observar en este caso. Miren que es una vista muy particular porque tiene dos cortes. Un corte transversal, que sería que nos estaría mostrando estas estructuras aquí: mesodermo paraaxial, aquí tenemos mesodermo intermedio, mesodermo lateral, un poco más allá. En la línea media tendríamos el tubo neural y la notocorda. Y en un corte paraaxial o parasagital, no sagital, porque si no estaría en medio del tubo neural, por eso es paraaxial, nos está mostrando los somitas.

En ese caso, el desarrollo, digamos, que esos estos somitas van a, van a tener un desarrollo bastante particular y van a tener una nomenclatura también bastante especial. Entonces, como podemos observar aquí, es que el somita, partiendo de que de que acá, digamos, en su porción, digamos, dorsal, pues vamos a tener la parte dorsal del somita, y que hacia el saco vitelino tendríamos la porción central del somita. Repetiría que la porción del saco estaría cerca de lo que es el ectodermo superficial. También podríamos decir que hay una parte que es medial y dentro de ese, dentro de ese somita, y esta es la que estaría cerca del tubo neural, que en este caso sería el ectodermo neural.

Entonces, va a, digamos, que en principio, por esta nomenclatura, entre nosotros decimos que el somita en su porción ventral y medial, serían estas células que están aquí de un color café un poco claro, van a generar lo que es el esclerotoma. Entonces, estas células mesenquimales del esclerotoma van a empezar a migrar, como las vemos aquí, y van a empezar a rodear lo que es el tubo neural. Siguen rodeando el tubo neural y a través de esto, lo que nos van a generar estos somitas, en este caso, las células centrales y mediales, va a ser el esclerotoma. Y este esclerotoma va a empezar a rodear el tubo neural para conformar las vértebras.

Entonces, aquí podríamos observar con claridad que tenemos el tubo neural, tenemos tubo neural. Allá ya podemos observar que están los esclerotomas. Y a su vez, partiendo de que se generan una serie de divisiones que van a estar, digamos, a una separación de células del esclerotoma en una porción caudal, en una porción craneal, y que está marcado por la migración de las células de la cresta neural y a su vez de axones que vienen desde el tubo neural, van a llegar a enervar estructuras particulares. Pero para el ejemplo del cual estamos hablando, pues nos van a separar esas células del esclerotoma.

Entonces, un cuerpo vertebral, como pudiéramos tener el que estamos observando aquí, va a estar conformado por la porción craneal del somita de abajo, sería este, y por la porción caudal del somita de arriba. Así es que esos dos se fusionan para producirnos un cuerpo vertebral. Pero a su vez, habría que aclarar que como tenemos del lado izquierdo y del lado derecho, un cuerpo vertebral va a quedar conformado por cuatro semitas, dos izquierdos y dos derechos. A su vez, cuando nosotros digamos que tenemos un proceso anómalo en este desarrollo de los somitas, su porción central y medial del esclerotoma y la conformación de estos cuerpos vertebrales, pues lo que podemos tener son fusiones de estas, de estos, de estos cuerpos vertebrales que se van a haber desarrollado mal, inclusive también de otras estructuras también óseas que están derivadas también de allí, de ese esclerotoma, que serían las costillas, y los pacientes tener tipo escoliosis, tipo escoliosis severa, en este caso en particular.

Entonces, ahora vamos a explicar que la porción dorsal, recuerden, cerca del ectodermo superficial y lateral, la vamos, digamos, a denominar dermatomiotoma. Si, entonces, ese dermatomiotoma nos va a producir la dermis. Recuerden que la epidermis está generada por el ectodermo superficial, mientras que la dermis es la que está debajo. 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